LaLonde was born in Richmond, California. Originally inspired by guitarists such as Frank Zappa, Snakefinger, East Bay Ray, Jerry Garcia, Jimi Hendrix, and his teacher Joe Satriani, LaLonde eventually became interested in early thrash metal bands such as Metallica and Slayer. LaLonde formed his first band in high school, a speed metal band called Blizzard. LaLonde left the band later on to join the first generation death metal band Possessed. Possessed’s landmark first album, Seven Churches, was released in 1985 when LaLonde was just 17 years old. Ler has been a member of Primus since 1989.